Issue was with our dear posebones again... when applying overrides we keep the same address/pointer for the IDs themselves, (which avoids us the need to remap their usages), but their inner data is often re-allocated. Therefore, we need once again to go over armature objects and invalidate their posebone pointers. This should also be back-ported to Blender LTS 2.83. Maniphest Tasks: T80078 Differential Revision: https://developer.blender.org/D8734