This issue occurred because versioning was applied twice.
The history:
* First I created 05aac73b45 which fixed the issue on main (4.5 at the time), version `405, 14`
* Then I created a backport for 4.4.3 f1e829a459 which also added versioning, version `404, 32`
That means all files created with 4.4.3 (`404, 32`) already had versioning
applied. However then opening it in 4.5 applied the versioning again since that only checked
if the version was at least `405, 14`
The fix is to exclude files that have already been saved with `404, 32`
Pull Request: https://projects.blender.org/blender/blender/pulls/142208